TOWARD A BETTER UNDERSTANDING OF TROPHIC DYNAMICS OF RIVER ECOSYSTEMS Michael D. Delong Large River Studies Center, Biology Department, Winona State University, Winona, MN 55987 Members of the Mississippi River Research Consortium have been considering the nature of trophic dynamics. Direct studies of riverine food webs are relatively new to both river science and the MRRC, but the history of the MRRC includes numerous studies that touch on the function of river ecosystems. The study of trophic dynamics dates to the second meeting of the MRRC, where Pahl and Varchman described the diet of three species of game fish. Studies linked to trophic processes appeared sporadically in the intervening years, including biomass and abundance of primary producers, bacterial production, and secondary production of benthic invertebrates. The late 1980s and early 1990s marked an effort to test the prevailing theory of riverine function of the time, the flood pulse concept. This is exemplified by examination of the FPC during and following the 1993 flood. Experimental studies added to our understanding of food webs and the interaction of a non- native species and temporal conditions. The development and application of other methodologies, such as stable isotopes, have expanded our understanding or riverine food webs further still. Application of stable isotopes of potential food sources and consumers brought the realization that the food web of the Upper Mississippi, like so many other large rivers world-wide, is supported by autochthonous production. Other techniques, such as fatty acids, have found their way into presentations at MRRC and the annual meeting will continue to see our understanding of riverine food webs expand as river scientists expand their study of the temporal and spatial dynamics of trophic processes. Keywords: history, food web, trophic, experiment, isotope, fatty acid, ecosystem theory